ԐzgtQh̍\Ƌ@\

<strong> VEcYE{ </strong>
iޗǐ[ȊwZpw@woCITCGX Aq`wj
emailF<a href="mailto:tsujih@bs.naist.jp"> V</a>
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̈Zr[, <strong>2</strong>, e004 (2013)@<a href="http://dx.doi.org/10.7875/leading.author.2.e004">DOI: 10.7875/leading.author.2.e004</a>
Hiroyuki Tsuji, Ken-ichiro Taoka & Ko Shimamoto: <strong>Structure and function of flowering hormone "florigen".</strong>

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<h2>v </h2>
@tQ͐Ảԉ蕪ÂqłC̐̂𖾂炩ɂ邱Ƃ͒AȊwɂdvȖ̂ЂƂłDߔN̕q`wIȉ͂̐iWɂCtQ̐̂FT/Hd3aƂ΂鋅^pNł邱Ƃ炩ɂȂCɍŋ߂ɂȂCtQê̓C@\̖{̂łtQ̂̌\̉𖾂ȂǁCqxł̗[܂ĂĂD܂CtQ͉ԂłȂWKČ`ÂȂǁCׂ@\Ƃ炩ɂȂĂDł́C킪E[hĂtQ̗jTςƂƂɁCtQ̍\Ƌ@\ɊւŐV̒mD

<h2>͂߂</h2>
@tQ́CAɉԉ`錈IȌʂƂāC1936Nɂ݂̑\ꂽ<a href="#R1"><sup>1)</sup></a>DtQ͔ԂƂ̂̂̎炷΂łȂCggDԂ̒R~jP[Vhgh{ƐBƂ̓]hƂdvȖ閣͓IȌΏۂƂȂC݂̑񏥂ĈȗCtQ̐̂𖾂炩ɂ邽ߐ̌ȂĂ<a href="#R2"><sup>2)</sup></a>DCw͂ɂ炸tQ̐̂͒̂܂܂łCg̐AzhƂ܂ł΂悤ɂȂ<a href="#R3"><sup>3)</sup></a>DߔÑfApq`wIȌ͂󋵂˔jC2007NCtQ̕q̂Hd3a/FTƂ΂^pNł邱Ƃ炩ɂ<a href="#R4"><sup>4,5)</sup></a>C2011NCtQ̎ê̓CсC̖{̂ƂȂ^pN̂̌\𖾂ꂽ<a href="#R6"><sup>6)</sup></a>i<a href="http://first.lifesciencedb.jp/archives/3509" target="_blank">V_r[</a> łfځjDł́CtQ̍\Ƌ@\ɊւŐV̒m܂Ƃ߁C񎦂tQ̓W]ɂċc_D
@͂߂ɁCtQ̊TOo܂ł̗jIȌo܂ɂĊTDA͔肵̂Cʏ͗tÂ邪CtHȂǓ̋G߂Ɨt̔~ĉԉ̌`iԐjJn<a href="#R3"><sup>3)</sup></a>Dōŏ̏dvȖ́CAv̂FċG߂𔻒fԐJnĂ邩łD1920NC^oR_CYɂĉԐ̃^C~O𒲍ʂ𑍍邱ƂɂCԐ̊Jn̍ő̌ƂȂ̂1̓̒ijł邱Ƃ炩ɂꂽ<a href="#R7"><sup>7)</sup></a>Dۂɂ萧䂳鐫́CʓIɌƂ΂DẢԐɂ萧䂳ԐĈȗC͓̋Gߐ̔ɐBsȂǂֈʉꂽ<a href="#R8"><sup>8)</sup></a>D݂ł́C̕q@\炩ɂC͊Tvƌ͌n̕Gȑݍpɂ萬Ă邱ƂmĂ<a href="#R8"><sup>8)</sup></a>D
@̏dvȖ́CAǂ̊튯ɂēFĂ邩łD܂܂Ȋ튯Ƃɂ튯ƂɓςꍇɁCԐ̃^C~O͂ǂω邩]ȂǂČvs튯͗tł邱Ƃ炩ɂȂ<a href="#R3"><sup>3)</sup></a>DCۂɉԂ̂튯́Ctł͂Ȃs̐[łDtŔFꂽ́Cǂ̂悤ɉԉ̂łs̐[ւƓ`B̂D1936NCVA̐AwChailakhyani`Cqj́CgtōČsֈړԉ𕪉镨h݂̑\CtQƖÂ<a href="#R1"><sup>1)</sup></a>i<a href="#F1">}1</a>jD̂̂Cgԉ𕪉ō͔|Ahgԉ𕪉Ȃō͔|AhւƐږ؂ƁC҂ɂԉ肪`ƂʂȂǂCtQ݂̑͋x悤ɂȂ<a href="#R3"><sup>3)</sup></a>DȗCtQ𕨎ƂĒP悤Ƃ͓IɍsꂽC̒Pɂ͂Ȃ<a href="#R2"><sup>2)</sup></a>D

<a name="F1"></a><div id="fig1-caption-text" style="display: none;"><strong>}1@tQɂԐ̐</strong>
A̗t͌͌nƊTv̏ƂɓvĂDԐɍœKȓFƁCtQR[h<em>FT</em>/<em>Hd3a</em>`q̔UDFT/Hd3a^pN͗ts܂ŒړCsɓBƉԉ̕JnDʐ^Hd3a-GFPZ^pN<em>Hd3a</em>`q̃v[^[ɂ蔭CľsiΐFHd3a-GFPZ^pŇuCԐF̓ClgD̎ƌujD<em>Hd3a</em>`q̃v[^[̓Cl̉Ԑ̗UłZɂėẗۊǑɓٓIɊD<em>Hd3a</em>`q̃v[^[ɂHd3a-GFPZ^pN𔭌ƁCtɂēٓIɔHd3asłώ@邱ƂłDiɂ́CtQɂԐ̐ɂ邨ȈqɂāC̓܂Ƃ߂D
<a href="http://leading.lifesciencedb.jp/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.1.jpg" target="_blank">[Download]</a></div>[hs_figure id=1&amp;image=/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.1.png&amp;caption=fig1-caption-text]

@tQ𓯒肵̓𐶗wIɖ炩ɂ悤Ƃ錤ɂāC킪̐Aw͐EĂDƂ1940NCsqȌԐATKIpƎ̌JnC̗ɍvĂDATKI̕i타TLł͎qtɑ΂1̒Z邾ŉԐ̗Uɏ\ȃtQƂCɁCԉ̐̕v邱ƂŃtQ̊ʓIɕ]łȂǁC̃tQ̐iɂ߂ėLȍޗłDCi߂ĂATKǏԐ̌̓tQ̈ړx̐͂߂ƂdvȔ炵<a href="#R3"><sup>3)</sup></a>C~ςCq悤ɁCtQ̕q̂̉𖾂ɂĂ킪̌҂vƂ̔wiɂȂ̂ȂD

<h2>1DtQ̐</h2>
@ߔÑClуVCkiYipq`wIȉ͂CtQ̕q̂Hd3a/FTƂ΂^pNł邱Ƃ炩ɂꂽ<a href="#R4"><sup>4,5)</sup></a>i<a href="#F1">}1</a>jDł́C̎ԉ𖾂̌o܂ɂĊTD
@1999NCsw̍r ̃O[v́ChCc̃O[vƂƂɁĈɃtQR[hĂ邱Ƃ炩ɂ<em>FLOWERING LOCUS T</em>i<em>FT</em>j`q𓯒肵<a href="#R9"><sup>9,10)</sup></a>DVCkiYỉԐ͒ɂđi邪C<em>ft</em>ψّ̂ɂĊSȉԐ̑i͋NȂDāC<em>FT</em>`q͗tɂtQ̍sɂe܂ŁCꂩ̃XebvɕK{ł邱Ƃ҂ꂽD<em>FT</em>`q̔镔ʂ͗ẗۊǑ⿕itȂǂɂh{̒ʂ蓹jłƂ<a href="#R11"><sup>11)</sup></a>CFT^pN͈ꌩƗtŋ@\Ăƍl悤D2005NɂȂC2̃O[v́CFT^pN̋@\镔ʂɂďdvȎ錋ʂ񍐂DFTSɋ@\ɂ́CsŔbZIP^̓]ʈqFDƂ̑ݍpKvłƂ񍐂ł<a href="#R12"><sup>12,13)</sup></a>D<em>FT</em>`q͂̔ʂtłɂ炸Cpʂsł\ꂽƂ悤D
@2007NɂȂCM҂̃O[vƃhCc̕ʂ̃O[v́CꂼCFT̃ClɂzOłHd3a^pNCсCVCkiYiFT^pNtQ̕q̂ł邱Ƃ錤ʂ񍐂<a href="#R4"><sup>4,5)</sup></a>D{eł́CM҂̃Clɂ<a href="#R5"><sup>5)</sup></a> 𒆐SɊT邪CVCkiYiɊւ񍐂ɂĂꂽǎ҂͌_<a href="#R4"><sup>4)</sup></a> QƂĂقD܂C<em>Hd3a</em>`q̃v[^[ڍׂɒC̃v[^[̓Cl̉ԐiZɂēٓIɗẗۊǑ⿕ɂĊCsł͂̃v[^[݂̊͂ȂƂꂽD<em>Hd3a</em>`qv[^[]ʂꐶmRNAtł̂݋~ς݂Csɂ͂قƂǒ~ςĂȂDŌɁCHd3a^pN̋Ǎ݂𒲂ׂ邽߁C<em>Hd3a</em>`qv[^[̐䉺ɂHd3a-GFPZ^pN𔭌CloCHd3a̋Ǎ݂GFP̌uwWɒǐՂꂽĎʁC]ʂmRNAقƂǂȂsɂāCHd3a-GFPZ^pŇuĂɌoꂽi<a href="#F1">}1</a>jD̂Ƃ́CHd3aẗۊǑ⿕sւƒړƂ<a href="#R5"><sup>5)</sup></a>DɁCHd3a-GFPZ^pN𔭌Cl͑炫ƂȂC܂C<em>Hd3a</em>`qƂ̃zOł<em>RFT1</em>`q̔RNAi@ɂmbN_EƉԐSɗ}ꂽƂCHd3aɂ̓Cl̉Ԑ𑣐i@\ƌ_ꂽ<a href="#R5"><sup>5,14)</sup></a>Ďʂ𑍍ƁCHd3a^pNړ̉ԐiVOigtQh̕q̂ł邱ƂꂽDľ_́Cr؂̃O[vɂVCkiYi̐ږ؎<a href="#R15"><sup>15)</sup></a>Cč̃O[vɂJ{`̐ږ؎<a href="#R16"><sup>16)</sup></a> ȂǂĂC݂ł́CtQ̎̂Hd3a/FTłƂ_͍L󂯓Ă<a href="#R17"><sup>17,18)</sup></a>D
@tQ̕q̂͒̈ړ^pNFT/Hd3ałDjIɂ͂̂ƂCtQ̓x点ł̂ȂDƂ΁CtQ̒o݂̎ɂāCΏۂ͂ق̐AzƓl̒ᕪqz肳ĂD̂߁CFT/Hd3a͐̉ߒŎꂽsꂽ肵ĂƐD܂CotQ̊mF@ƂāCoɐÂȂǂZЂ͓hz荪z킹肵ĉԐώ@@ƂĂD̏ꍇCFT/Hd3a敪ɑ݂ĂƂĂq20,000Ƃqł邽ߍזE𓧉߂łCāCmF邱Ƃ͍ł낤D

<h2>2DtQ̐wIȓ</h2>
@炳́CŐV̌ʂ疾炩ɂȂCtQ̊{̂ł^pŃCсCtQ̍זEɂéCɁC̔WJtQ̐V@\ɂĊTDoꂷ邨ȈqɂĂ܂Ƃ߂̂ŁCQƂ̂ǂݐiłقi<a href="#F1">}1</a>jD
@FT/Hd3a̓zXt@`WG^m[A~^pNiphosphatidylethanolamine binding proteinFPEBPjƂ悭1\^pNłDzXt@`WG^m[A~^pN̓̈łzXt@`WG^m[A~ƌn̃^pNƂăEV̔]݂Ĉ̂C̐ɕՓIɑ݂邱Ƃ炩ɂȂDq20,000̏ȋ^pNŁCzXt@`WG^m[A~̃_̕AjI|Pbg̈Ƃ΂鏬Ȃڂ݂DtQ3\ɂẮChCc̃O[vFT <a href="#R19"><sup>19)</sup></a>CM҂̃O[vHd3a <a href="#R6"><sup>6)</sup></a> ɂČ\񍐂Ăi<a href="#F2">}2</a>jD2݂͌ɔɎ\CAjI|Pbg̈̍\ۑĂ邪CzXt@`WG^m[A~^pNɂ郊_̔Fɏdvȃ`Vc͕ۑĂȂD݂̂ƂCtQ̉Ԑi@\ƃzXt@`WG^m[A~Ƃ̊֘A͕słD

<a name="F2"></a><div id="fig2-caption-text" style="display: none;"><strong>}2@tQ̍\</strong>
iajCl̃tQłHd3ǎ\DzXt@`WG^m[A~^pN̓łAjI|Pbg̈ƁC̓t߂ɃtQɓITyr87iΐFj݂D14-3-3Ƃ̌̈ԐFŁCZOgB̈FŎD
ibjVCkiYĩtQłFTƉԐvbT[TFL1̌\D
icjFTTFL1Ƃ̏dˍ킹D2̑S̍\͂悭Ă邪CZOgB̈͑傫قȂĂD
<a href="http://leading.lifesciencedb.jp/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.2.jpg" target="_blank">[Download]</a></div>[hs_figure id=2&amp;image=/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.2.png&amp;caption=fig2-caption-text]


<h2>3DtQƑݍp^pN̔</h2>
@܂łɒmĂzXt@`WG^m[A~^pN̐wIȋ@\́CtQɂԐ̑i̕q@\𐄑邱Ƃ͍łD̂悤ȏꍇCΏۂł@\s̃^pNǂ̂悤ȃ^pNƑݍpĂ邩m邱Ƃŋ@\̐ɂďdvȎ肪肪邱ƂDɏqׂ<em>FD</em>`q́CԐxψّ̂ƂĂłɕ񍐂Ă<em>fd</em>ψّ̂̌`qłD<em>FD</em>`qbZIP^̓]ʈqR[hĂCFTFD̗҂̉ߏ蔭ɂԉ蕪gD`q̂ЂƂł<em>AP1</em>`qُ̔IɗUł<a href="#R13"><sup>13)</sup></a>DāCyc[nCubh@ɂFTFDƂ̑ݍpoꂽ<a href="#R12"><sup>12,13)</sup></a>D̑ݍpɂ́CFDC[ɑ݂CCa<sup>2+</sup>ˑveCLi[[icalcium-dependent protein kinaseFCDPKjɂ胊_Ɨ\zA~m_z񃂃`[tThr-Ala-Prȏ݂K{ł<a href="#R12"><sup>12)</sup></a>DāC̃`[tFD͉Ԑɂ@\rDȏォCFTFDƂݍp邱ƂFTɂԐ̑iɏdvłƍlꂽ<a href="#R12"><sup>12)</sup></a>ĎʂC]ʈqFD́Cꎩ̂ɓ]ʊ\͂ȂCFTƌ邱Ƃɂ]ʊ\lWI`q̓]ʂCƂfoꂽ<a href="#R19"><sup>19)</sup></a>D
@M҂̃O[vCtQ̋@\̗̂ߍyc[nCubh@ɂHd3aݍp^pN̒Ts<a href="#R6"><sup>6)</sup></a>ĎʁC14-3-3^pN̂ЂƂłGF14cCVCkiYiFD̃ClɂzOłOsFD1ȂǂHd3aƑݍp^pNƂēꂽi<a href="#F3">}3a</a>jD14-3-3R[h`q̓Cl̃Qmɂ8݂ĂCꂼ̎YGF14aGF14ĥ̖Ă邪CHd3aƌ̉\Ȃ̂͂̂4CGF14bCGF14cCGF14eCGF14fłD4̋@\̓tQ̋@\ɊւĖ{Iɓłƍlꂽ̂ŁCł͂܂Ƃ߂14-3-3ƂԁD̃A~m_zrƂC14-3-3Hd3aݍp^pNC[ɂ́CɏqׂFD-FTݍp`[t<a href="#R12"><sup>12)</sup></a> ɗގSAP`[tSer-Ala-ProʂČoC͍yc[nCubh@ɂHd3aƂ̑ݍpɕK{łDCxɐg^pNp<em>in vitro</em>ɂ鑊ݍpł́CHd3a14-3-3Ƃ̂̑ݍp͌oꂽCHd3aOsFD1Ƃ̂̑ݍp͌ołȂD̎ʂƁC14-3-3̌`[tł郂[h1SAP`[tĂ邱ƁCɁC14-3-3SAP`[t͂ȂƂCHd3aƂ̑ݍp^pŃC14-3-3ԐړIɑݍpĂƍli<a href="#F3">}3a</a>jDyc[nCubh@ɂĊώ@ꂽHd3aƂ̑ݍp^pNƂ̌́Coyɓ݂14-3-3ԐړIȂ̂łƐłDہCOsFD114-3-3Ƃ<em>in vitro</em>̎nɂĂݍpCɁCψى͂ɂSAP`[tSer̃_OsFD114-3-3Ƃ̑ݍpɏdvł邱Ƃ킩Dyc[nCubh@̓^pN̑ݍpȕւɎł邷ꂽnł邪Coyɓ݂^pNɂԐړIȑݍpo\ЂłD܂Cv_EAbZCȂ<em>in vitro</em>ɂ鑊ݍp͂ɂĂCp^pN̐x\ɍȂ΃^pN̎YɗpזÊ݂ݍpɉeꍇ邽߁Cʂ̉߂ɂ͒ӂKvłDFTFDƂ̑ݍp̉͂̌ʂ́Ĉ悢PƂ邩ȂD

<a name="F3"></a><div id="fig3-caption-text" style="display: none;"><strong>}3@tQ̂̍\</strong>
iajHd3aCGF14COsFD1̖͎}Dꂼ̑ݍp̈DNLSFjǍ݃VOiCNESFjOڍsVOiD
ibjDNÃtQ̂̍\fDZOgBAjI|Pbg̈͊OʂɘIoĂD
icjHd3a14-3-3Ƃ̑ݍpʁi㕔jƁC14-3-3OsFD1Ƃ̑ݍpʁijDHd3aOsFD1̂ɒړIȌ݂͂ȂD
<a href="http://leading.lifesciencedb.jp/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.3.jpg" target="_blank">[Download]</a></div>[hs_figure id=3&amp;image=/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.3.png&amp;caption=fig3-caption-text]


<h2>4DtQ̂̍\</h2>
@M҂̃O[v́CHd3aC14-3-3COsFD1̑ݍp̏ڍׂ𒲂ׂ邽߁C3̃^pNȂ^pN̗̂̍\͂sC\2.4𕪉\Ō肷邱Ƃɐ<a href="#R6"><sup>6)</sup></a>iPDB IDF<a href="http://service.pdbj.org/mine/Detail2?PDBID=3axy&PAGEID=Summary" target="_blank">3AXY</a>jDɂCOsFD1ƂĂ14-3-3Ƃ̌ɕKv\ȃ_Ser192܂C[9A~m_cȂfЂpDꂽ̂̍\́CHd3aC14-3-3COsFD1ꂼ2qȂW^̃weZʑ̂`Ăi<a href="#F3">}3b</a>jDāCʑ̂`14-3-3W̒ɂ邭ڂ݂ɁC_ꂽOsFD1͂܂荞݁C̏㑤Hd3a1qEΏ̂ɗČĂDHd3aOsFD1Ƃ̂ɒړIȑݍp݂͂Ȃi<a href="#F3">}3c</a>jD̂̕tQ́iflorigen activation complexFFACjƖÂD
@14-3-3͐^jɍLۑĂ镪qʖ30,000̎_^pNłD14-3-3ƂÓCEV̔]ɑ݂^pNԗIɕނۂɁCN}gOtB[ɂ敪ԍ14ƓdCjł̈ړʒu3.3ɑ݂^pN14-3-3ƖƂɗRD14-3-3ɂ͕̈قȂ镪q킪݂ĈŃzʑ̂邢̓weʑ̂`D14-3-3ʑ̂̓_Zc邢̓_XIjcFĂ܂܂ȃ^pNƌC̋@\𐧌䂷D14-3-3ƌ^pN̑ɂ́C14-3-3ɂF郊_Zc邢̓XIjc̎͂ɁCʂA~m_z񃂃`[tł郂[h1iArg-X-X-pSer-X-Proj邢̓[h2iR-X-X-X-pSer-X-ProjoD
@tQ̂̏ꍇC14-3-3OsFD1Ƃ̌SAP`[t̃_Zc̔FɂC14-3-3ɓT^Iɂ݂錋łDɑ΂āC14-3-3Hd3aƂ̌̓_ɔˑIȐVKȌłDāČlC14-3-3OsFD1Hd3a̗ɓɌ邱Ƃ\ɂĂDtQ̂\3̃^pN̑ݍpʂڍׂɂ݂ƂCOsFD114-3-3Ƃ̑ݍpɂẮCOsFD1̃_ꂽSer19214-3-3̉̃_yv`h|Pbgɂ͂܂荞݁CɁCSAP`[t̑ŜFĂi<a href="#F3">}3c</a>jDC14-3-3Hd3aƂ̑ݍṕCHd3a̒[v̈ɑ݂2̓˂oAMjcArg64Arg13214-3-3̏㕔ɂ_̂ڂ݂ɂ͂܂荞݁CɁCHd3a̖{̂14-3-3C[̂փbNX̂ɂa̍aƍLݍpĂi<a href="#F3">}3c</a>jDCl̃tQ̂ɂĊώ@ꂽHd3a14-3-3C14-3-3OsFD1̑ݍpʂ́CƂɃClȊO̍AFTC14-3-3CFDɂĂxɕۑĂ<a href="#R6"><sup>6)</sup></a>DVCkiYiɂāCFT14-3-3Ƃ̑ݍpłɕ񍐂Ă<a href="#R20"><sup>20)</sup></a>DȏォCtQC14-3-3C]ʈqFDɂ镡̂̌`͍ẢԐ̐ɂċʂđ݂CtQׂĂ̐Aɋʂɂ͂炭qՂɂȂĂƍlꂽD
@A̍זEɂāCtQ̂̓Cl<em>OsMADS15</em>`qVCkiYi<em>AP1</em>`qȂǉԉ``q̃v[^[̈Ɍ]ʂĂƍl<a href="#R6"><sup>6,13)</sup></a>DVCkiYi<em>AP1</em>`qv[^[̉͂́CC{bNXz܂ޓ̗̈FTFDɑ΂鉞\邱Ƃ񍐂Ă<a href="#R6"><sup>6,13)</sup></a>D܂C<em>in vitro</em>ɂăCl̃tQ̂C{bNXzƂ邱ƂQVtgAbZCɂ薾炩ɂĂ<a href="#R13"><sup>13)</sup></a>DSFDƕWIƂȂDNA܂ރtQ̂̌\̗͂͂܂ȂCmł铮bZIP^̓]ʈqCREBDNAƂ̂̍̕\Ƃɍ쐻ꂽtQ̂DNAƂ̂̃̕f\i<a href="#F3">}3b</a>jł́CHd3aOsFD1Ƃ͒ڂɌ邱ƂȂCHd3a14-3-3OsFD1DNAɈɕێ悤ʒuĂDāCHd3a̓tQ̂̈艻ɂ^Ă\lꂽD

<h2>5DtQ̂̌`@\FtQêƂĂ14-3-3</h2>
@Cl̃vgvXgpătQ̂זEɂč\z@\ڍׂɒׂꂽ<a href="#R6"><sup>6)</sup></a>DGFPȂǂ̌u^pNƂ̗ZɂtQ̂̌X̍\^pN̍זEł̋Ǎ݂ώ@ꍇCHd3a͊jƍזEɁC14-3-3͂قƂǂזEɁCOsFD1͊ĵ݂ɋǍ݂ĂCtQ̃fɂĕ̂`͂3̃^pN̋Ǎ݂͂ꂼقȂĂi<a href="#F4">}4a</a>jDɁĈ̍זEɂǍ݂񕪎quԁibimolecular fluorescence complementationFBiFCj@ɂ蒲ׂꂽDBiFC@́Cݍp\𒲂ׂ悤Ƃ2̃^pN̂ꂼ2u^pNZC2̃^pNݍpƌu^pN̗̍\č\Čuώ@CƂɂ@łDBiFC@ɂHd3a-14-3-3̂̌`͍זEɂČoꂽi<a href="#F4">}4a</a>jDƂ낪CɂCFP-OsFD1Z^pNƁCHd3a-14-3-3̂OsFD1ɈˑIɍזEjւƋǍ݂ςC3̃^pNjɏW܂邱Ƃώ@ꂽi<a href="#F4">}4a</a>jDāCBiFC@ɂĂCHd3a-OsFD1̂͊jɂČoꂽD܂CVCkiYiɂĂFT-FD̂BiFCVOi͊jɂČoDl킹ƁC14-3-3͍זEɂHd3aƍŏɌtQêƂċ@\CHd3a-14-3-3̂͊jւƈڍsOsFD1Ƒݍpƍlꂽi<a href="#F4">}4b</a>jD

<a name="F4"></a><div id="fig4-caption-text" style="display: none;"><strong>}4@tQɂԐ̐</strong>
iajvgvXgɂזEł̑ݍpDHd3a14-3-3ƂBiFCݍp͂ɍזEɂĊώ@邪COsFD1Hd3a-14-3-3̂͊jɋǍ݂悤ɂȂD
ibjf}DۊǑ⿊ǂƂČsւƉ^΂ꂽtQ́CsgDזE̍זEɂĎêł14-3-3ƌD̃tQ14-3-3Ȃ镡̂́Cɓ]ʈqłFDƌătQ̂`Cjɂ<em>AP1</em>`q<em>OsMADS15</em>`qȂǉԉ`ɂ`q̓]ʂ邱ƂŁCԐЂNDPF_D
<a href="http://leading.lifesciencedb.jp/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.4.jpg" target="_blank">[Download]</a></div>[hs_figure id=4&amp;image=/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.4.png&amp;caption=fig4-caption-text]

@܂ŏЉǂ̎@CtQ̂\^pN2qԂ̑ݍp𖾂炩ɂ̂łC3qgɁhЂƂ̂̕ɑ݂邱Ƃ̂ł͂ȂDM҂̃O[v́CBiFC@ƌuGlM[]/uifluorescence resonance energy transfer/fluorescent lifetime imagingFFRET/FLIMj@Ƃgݍ킹@ɂC̓_𖾂炩ɂ邱Ƃ݂DFRET/FLIM@́C2q̂̌uGlM[ړiFRETjɂu̕ωϑ邱ƂɂC2qߐڂĂ邩ǂʓIɕ]@łD̓Iɂ́CHd3a14-3-3Ƃ̑ݍpBiFC@ɂ茟oCɁCBiFC@ɂčč\ꂽu^pNVenusCFP-OsFD1Z^pNƂ̑ݍpFRET/FLIM@ɂ茟o邱ƂŁC3̃^pNȂ镡̂̌o݂D͂̌ʂ́CזEɂĂ3̃^pNȂtQ̂͊jɂČ`Ă邱Ƃ<a href="#R6"><sup>6)</sup></a>D
@fi<a href="#F3">}3b</a>jł́CזEɂČ`ꂽHd3a-14-3-3̂ɂĊjɑ݂OsFD1Ƒݍp̂C܂łĂȂDɂẲ͍͉ۑł邪Ĉ݂ƂC̂悤ȉL͂łDȂ킿CזEɂĖ|󂳂ꂽOsFD1זEHd3a-14-3-3̂ƑݍpC`ꂽtQ̂OsFD1̂jǍ݃VOî͂炫ɂjւƈڍsCƂ̂ł<a href="#R21"><sup>21)</sup></a>D̉T|[gmƂāC14-3-3̂Hd3ä14-3-3̊jOڍsVOiƏdȂĂCHd3aƂ̌ɂ肻̋@\͑jQƗ\z邱ƂDЂƂCŋ߂ɂȂ蓾ꂽOsFD2ɂĂ̒m<a href="#R21"><sup>21)</sup></a>DOsFD2͗t̔Ɋ֗^bZIP^̓]ʈqŁCSAP`[tގz14-3-3ƌCHd3aƃtQ̂`邱Ƃł<a href="#R21"><sup>21)</sup></a>DOsFD1ƔrƊjǍ݃VOi̔zɑ傫ȈႢ݂C炭̂߂ɁCjւ̋Ǎ݂̌OsFD1ɔגႢ<a href="#R21"><sup>21)</sup></a>DCSAP`[tގzɕψق𓱓14-3-3Ƃ̌\rCHd3aătQ̂`肷ƁCOsFD2̊jւ̋Ǎ݂̌͒㏸<a href="#R21"><sup>21)</sup></a>D̊ώ@ʂ́Cɏqׂɂ肤܂ł̂łD

<h2>6DtQ̂̋@\</h2>
@Hd3ả̕WI`qƂ<em>OsMADS15</em>`q񍐂Ă<a href="#R14"><sup>14)</sup></a>D<em>OsMADS15</em>`qMADS{bNX`qt@~[̂<em>AP1</em>`qTut@~[ɑDVCkiYiɂāC<em>AP1</em>`q͉Ԋ튯̌`ABCfɂăNXA`qƂċ@\邾łȂCԐɂĉԉ蕪gD`qƂĂ@\CFTFDɂ萳ɐ䂳邱Ƃ炩ɂĂ<a href="#R12"><sup>12,13)</sup></a>Dŋ߂ɂȂCCl<em>AP1</em>`qTut@~[3̃o[C<em>OsMADS14</em>`qC<em>OsMADS15</em>`qC<em>OsMADS18</em>`qƁC<em>SEP</em>`qTut@~[̃o[ł<em>PAP2</em>`qi<em>OsMADS34</em>`qj̎ldψّ̂ł́CBւ̑]NȂƂꂽ<a href="#R22"><sup>22)</sup></a>DŁC<em>OsMADS15</em>`q̊Ԑ̃}[J[ƂāCԐɂHd3aC14-3-3COsFD13҂̑ݍp̏dvڍׂɒׂꂽ<a href="#R6"><sup>6)</sup></a>DߓIɂāC<em>OsMADS15</em> mRNA̗ʂ<em>Hd3a</em>`q<em>OsFD1</em>`qƂɔꍇɂ̂ݑDC14-3-3Ƃ̑ݍpψق𓱓<em>Hd3a</em>`qC邢́C14-3-3Ƃ̑ݍpψق𓱓<em>OsFD1</em>𔭌ꍇɂ́C<em>OsMADS15</em> mRNA̔̏㏸͂قƂǊώ@ȂDŒӂׂ_́CvgvXgɂ͓ݐ14-3-3ʂɔĂ邱ƂłD̂߁C14-3-3̈ߓIȔ̗L<em>OsMADS15</em> mRNA̔㏸ɂقƂǉeȂDŁCCl̃QmHd3aƌ̉\14-3-3R[h`q4݂邪Cɑ΂RNAixN^[𓱓邱ƂɂHd3aƌ̉\14-3-3ׂ̔ămbN_EƂC<em>Hd3a</em>`q<em>OsFD1</em>̔Ɉˑ<em>OsMADS15</em>`q̊͗}ꂽDȏ̎ʂ́CtQ̂̌`͉̕WI`q̊ɕK{ł邱ƂD
@ɁČ`]ClpCtQ̂̌`͎ۂɉԉ̌`ɂĕKvł邱Ƃ炩ɂȂ<a href="#R6"><sup>6)</sup></a>D<em>Hd3a</em>`qߏ蔭`]Clł́C̏o͌ɑiꂽ<a href="#R5"><sup>5,6)</sup></a>C14-3-3Ƃ̑ݍpψ<em>Hd3a</em>`q̉ߏ蔭ł͂̉Ԑ̑i@\͎ꂽ<a href="#R6"><sup>6)</sup></a>D܂C<em>OsFD1</em>̔RNAi@ɂmbN_EƁCVCkiYi<em>fd</em>ψّ<a href="#R12"><sup>12,13)</sup></a> gERV<em>dlf1</em>ψّ<a href="#R23"><sup>23)</sup></a> ƓlɁCԐ͒xDāC<em>OsFD1</em>ߏ蔭ƉԐ͑iꂽDC<em>OsFD1</em>̉ߏ蔭ɂԐ̑iʂ́C14-3-3ƍPIɌ̉\ȃ_͕ψق𓱓<em>OsFD1</em>ɂ̂݌łD̂Ƃ́C<em>OsFD1</em>̉Ԑ̐ւ̊֗^ƂƂɁCOsFD1̃_̐dvȐv̂ЂƂł邱ƂD
@ԉ``q̊ɂtQ̂̏dv͖炩ɂȂCtQ̂ǂ̂悤ȕq@\ɂ艺̕WI`q̓]ʂĂ̂͂܂炩ɂĂȂDOsFD1/FDbZIP^̓]ʈqł邪CPƂł͉̕WI`qȂD܂COsFD1Hd3a/FTɂ͓]ʈqɓT^Iɂ݂_̈̂悤ȃhC݂͂ȂDFT̏ڍׂȃhC͂́CZOgBƂ΂郋[v̈ƃAjI|Pbg̈悪Ԑ̑i@\ɏdvł邱Ƃ炩ɂĂ<a href="#R19"><sup>19,24)</sup></a>D̗̈́CtQ̂̍\fł͂̕\ʂɘIoĂ邱Ƃi<a href="#F3">}3b</a>jC̗̈ɖ̓]ʊqN[g邱Ƃɂ艺̕WI`q̓]ʊЂN̂ł͂ȂƗ\zĂ<a href="#R6"><sup>6)</sup></a>D

<h2>7DtQ̑lȋ@\ƃtQ</h2>
@͐Â܂܂ȐۂɂĂ邱ƂÂmĂDWKC̉siCǰ`ZŗU錻ۂ͂̑\IȗłDš`ɂĂCtɂč鉼z̉s`zi`[xQj`UƂ񏥂ꂽ<a href="#R25"><sup>25)</sup></a>Dł́C̂悤ȎC`[xQtQƓ̕ł\ɂĕ񍐂ꂽi<a href="#F5">}5a</a>jDԐɂ̈قȂ^oRiiZCCjƃWKCƂڂ؂C؂łWKCɂš`𒲂ׂƂC`ώ@ꂽ̂͂̐ڂ؎̏ꍇɂĂ؂ł^oR̉ԐUĂƂ݂̂ł<a href="#R25"><sup>25)</sup></a>D̂Ƃ́CقȂAłꂽԐU镨ڂ؂ɂ`BCWKCɂĉš`Uł邱ƂD

<a name="F5"></a><div id="fig5-caption-text" style="display: none;"><strong>}5@tQ̑@\ƃtQ̍\^pNf</strong>
iajWKCɂڂ؂ɂš`̗UDZɂš`UWKCƁCɂԐU^oRƂڂ؂CɂĐ炳ƁC^oRł͉ԐCWKCł͉š`UD
ibj]ʈqfDtQns̐[ւƗACŎêł14-3-3ƕ̂`Cɉs`]ʈqƃtQ̗l̂̕`ƁCs``q̓]ʂs`D
icjtQƉԐvbT[fDtQ̂̂Ȃ̃tQԐvbT[TFL1/RCNɒuԐ}̂́Cԉ``q̓]ʂ}Ă邪CtQTFL1/RCNƂuƃtQ̂ƂȂCԉ``q̓]ʂ𑣐iԐUD
PF_D
<a href="http://leading.lifesciencedb.jp/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.5.jpg" target="_blank">[Download]</a></div>[hs_figure id=5&amp;image=/wordpress/wp-content/uploads/2013/03/Shimamoto-2.e004-Fig.5.png&amp;caption=fig5-caption-text]

@M҂͍ŋ߂ɂȂCCl̃tQR[h<em>Hd3a</em>`qWKCɂĉߏ蔭ƁC{͉š`UȂɂĂs`邱Ƃo<a href="#R26"><sup>26)</sup></a>D̉ߏ蔭̌ʂ͐ڂ؂̓`BƂCtQ`[xQƂĂ@\Ă邱ƂꂽDŋ߂̌Ĉ悤ȉԐ̐̂ق̃tQ̋@\炩ɂ͂߂ĂDƁC|v̒Zɂ鐬̒~Ƌx̌`<a href="#R27"><sup>27)</sup></a>CeTCiTgE_CRjɂԐ̗}<a href="#R28"><sup>28)</sup></a>Cg}ggERVɂ`Ԍ`<a href="#R29"><sup>29,30)</sup></a>CVCkiYiɂCEJ̐<a href="#R31"><sup>31)</sup></a>Cg}gɂʎ<a href="#R32"><sup>32)</sup></a> ɂāC<em>FT</em>`q̃zÔĂ邱Ƃ񍐂ĂD|v̏ꍇC2̈قȂ<em>FT</em>`qzOC<em>PtFT1</em>`q<em>PtFT2</em>`q𕪒Sĉh{ѐB𐧌䂵Ă<a href="#R27"><sup>27)</sup></a>D<em>PtFT1</em>`q͓~̂ɔB̊Jňɂ̂ɑ΂C<em>PtFT2</em>`q͉h{ɔh{x̋x̑jQ𑣐iƍlĂDeTC̏ꍇC2̈قȂ<em>FT</em>`qzÔC<em>BvFT2</em>`q͉Ԑ̑iɕK{ł̂ɑ΂C<em>BvFT1</em>`q͉Ԑ̗}ɂ͂炫C̔̌tɂԐɏdvł<a href="#R28"><sup>28)</sup></a>Dg}g̏ꍇC<em>FT</em>`q̃zOł<em>SFT</em>`q́CԐ݂̂Ȃ炸ť`Ԃ܂߂CʓIȌ`Ԍ``qƂċ@\Ă<a href="#R29"><sup>29)</sup></a>DɁC̈`qʂʎʂɂdvł邱Ƃ񍐂Ă<a href="#R32"><sup>32)</sup></a>Dŏdvȓ_́CHd3aɂ14-3-3Ƃ̌ɂA~m_ćCɂǂFTzOɂĂxɕۑĂƂłDāCFTzO14-3-3ƌłƍlDЂƂ̖͓Iȉ́CCl̃tQ̂ɂ]ʈqOsFD1C14-3-3Ƒݍpłʂ̓]ʈqɒuŃtQ̗l̂̕`Cԉ`֘A`qƂ͈قȂWI`q̔䂪ȂĂCƂ̂łi<a href="#F5">}5b</a>jD̂悤ȓ]ʈq̌ƂāCtQ̂̌`\Ԑł͂Ȃť`ɂƍlOsFD2悤<a href="#R21"><sup>21)</sup></a>D
@[̂́C@\IɝhR<em>FT</em>`q̃zOɂĂ͍\ɂႢ݂邱ƂłDʂɁCԐ̑iɂ͂炭FTzÕA~m_z͔ɍxɕۑĂ邪CPtFT2BvFT1̂ZOgB̈ɂ͈ʓIȃRZTXzƔrăA~m_z̒u݂DZOgB̈́Cɏqׂ悤ɁCtQɂ鉺`q̓]ʊɏdvȗ̈łDBvFT1̃ZOgB̈ɂ݂A~m_ɒu̓ꂽψBvFT2́CԐiԐ}ւƋ@\]<a href="#R27"><sup>27)</sup></a>DɂẮCZOgB̈ႢɂقȂ]ʊq邱Ƃŋ@\̕ϊN\l悤D

<h2>8DԐvbT[TFL1ƃtQ</h2>
@ԐvbT[`q<em>TERMINAL FLOWER1</em>i<em>TFL1</em>j́C炫Ɖԏ̗L̕\^VCkiYiψّ̂瓯肳ꂽ<a href="#R33"><sup>33)</sup></a>D<em>tfl1</em>ψّ̂͑炫ƂȂCԏgDԂɕϊ邽ߒ[ɉԂ`LԏƂȂ邪C<em>TFL1</em>`qߏ蔭Ƌtɒx炫ƂȂ薳悤ɂȂDM҂̃O[vC<em>TFL1</em>`q̃ClɂzOł<em>RCN</em>`q̉ߏ蔭Ԑ̗}ɂ͂炭Ƃ񍐂Ă<a href="#R34"><sup>34)</sup></a>D܂ŋ߂ɂȂāCNẢԐ̗}<a href="#R35"><sup>35)</sup></a> oC`Sɂ݂lG炫<a href="#R36"><sup>36)</sup></a> ̌`qƂ<em>TFL1</em>`q̃zO񍐂ĂCJԂ̋Gߐɂ<em>TFL1</em>`qzȌdv炩ɂĂĂD̒ḿC<em>TFL1</em>/<em>RCN</em>`qԐɊւ<em>FT</em>/<em>Hd3a</em>`qƂ͐΂ɂ͂炭ƂĂDƂ낪CTFL1/RCNFT/Hd3aƓlɃzXt@`WG^m[A~^pNt@~[ɑ邱ƂCFT/Hd3aTFL1/RCN͉Ԑ̐̋ʂoHɂĝhRIɂ͂炭ƍlĂ<a href="#R9"><sup>9,10,19,24)</sup></a>D
@<em>FT</em>`q<em>TFL1</em>`qƂ̝hRIȋ@\̕ɂẮC̎Ył^pN̍\ƂɂIȉ͂ȂĂDAjI|Pbg̈̎ӂɈʒuFTTyr85iHd3ałTyr87jFTTut@~[ɂāCTFL1His88TFL1Tut@~[ɂāCSɕۑĂ邱ƂC̃A~m_c̒usꂽ<a href="#R24"><sup>24)</sup></a>ĎʁCFTTyr85HisɒuFTߏ蔭̂Ɣגx炫ɂȂDC<em>FT</em>`q<em>TFL1</em>`q̃L`q̉Ԑɑ΂ʂԗIɒׂCZOgB̈悪<em>FT</em>`q<em>TFL1</em>`q̂Ɠِ炷Ƃ炩ɂꂽ<a href="#R19"><sup>19)</sup></a>DZOgB̈FTTFL1̌\rƂ傫قȂĂ郋[v̈ɑĎʂ́CAjI|Pbg̈ZOgB̈̈ႢFTTFL1Ƃ̋@\̕ɏdvł邱Ƃ<a href="#R19"><sup>19,24)</sup></a>D
@ŏdvȓ_́CHd3aɂ14-3-3Ƃ̌ɂA~m_cTFL1/RCNTut@~[ɂĂxɕۑĂ_ł<a href="#R6"><sup>6)</sup></a>DہCVCkiYig}gTFL1zO14-3-3Ƒݍp邱Ƃłɕ񍐂Ă<a href="#R20"><sup>20)</sup></a>DTFL1/RCNɂԐ̗}̕q@\ɂāCԐ̑iɂtQ̃fƂɂ̂悤ȉlDȂ킿Ch{ɂ̓tQ̂̃tQ̈ʒuTFL1/RCN荞񂾉Ԑ}́iflowering repression complexFFRCj`ԉ``q̓]ʂ}Ă邪CtQsɓBƁCԐ}̂TFL1/RCNtQƒuătQ̂ւƓ]Cԉ``q̓]ʂiԐUi<a href="#F5">}5c</a>jD܂CsgDɂTFL1/RCN]ʈq14-3-3Ƃ̂̕tQƋ邱ƂɂtQ̊̃oX𐧌䂵Cێ̂ɂĂ\li<a href="#F5">}5c</a>jD̃fɂāCtQ̋@\̗}ɂTFL1/RCN14-3-3ƌ邱ƂɂtQtQ̂`̂jQ邾ł\łƍl邪CԐ}̂ɂTFL1/RCÑAjI|Pbg̈ZOgB̈ɓ]ʐRvbT[ĉԐ`q̓]ʂϋɓIɗ}Ă\<a href="#R37"><sup>37)</sup></a>D

<h2></h2>
@܂łɓꂽm𑍍ƁCtQłHd3a/FT͗tōꂽ̂CۊǑƂČs܂ŒړCsזE̍זEɂ14-3-3ɎeHd3a/FT-14-3-3̂`D̂̂CHd3a/FT-14-3-3̂͊jւƈڍsCFDƂɍ̂̕łtQ̂`ĉԉ``q̓]ʂCԐJnƍlDFT͒PȂgԐhzɂƂǂ܂炸CWKCɂš`܂ނ܂܂Ȍ`Ԍ`𐧌䂷Cړ̑@\złƔF悤ɂȂĂD̑@\SqՂƂāCtQ̂̍\^pŇflD
@tQ̗i߂邤ł́C̂悤ȏdvȉۑ肪cĂDЂƂ́CtQ̂Ɍ]ʊq̓CDNA܂߂SȃtQ̗̂̍\̌łDɂCtQ̐wIȖtQ̂̌`K肷\Iȗv炩ɂȂ邾낤D̂̉ۑ́CtQ̑@\̕q@\𖾂炩ɂ邱ƂłD܂̓tQ̃f\z悤ȓ]ʈq̒T}ł낤DtQ̒ړɂ镪q@\̗dvłDŋ߂ɂȂCFT⿊ǔזE⿕vfւ̈ڍsɂ^pNƂFTIP1ƂC2hC^pN񍐂ꂽ<a href="#R38"><sup>38)</sup></a>DCFTIP1ɂFTA̕q@\͖炩ɂĂȂDɁC⿕vfsgD̍זEւ̗A@\͕ŝ܂܂łD܂CtQɂԐ̐̍ۂɌsgDɂĂǂ̂悤ȕωЂNĂ̂CȂ킿CsgDɂ銲זE̐̓]ߒ̑S̑𗝉KvDɂ́CGsQm⏬qRNACgXNvg[ω̖ԗIȉ͂L͂Ȑ؂ƂȂ邾낤D܂CtQ̐זEC[WOƔ͂Ƃgݍ킹邱ƂɂCtQɂsgD̋ԓIȐ̗iނ낤D
@tQ̌͊bȊwɂƂǂ܂ȂDẢԐlדIɐ䂷Zp͔_ƂɂĔɗLpłDCgԍ炩ꂳh̘bɂ悤ɁCtQAɂӂ肩ł͉NȂ낤DȂȂCtQ̎ê͍זEɂ邪CtQ͍̂זEʉ߂łȂ^pNłDtQ̓^ɂԐ̐ɂ͐A̍זEɃtQ荞܂@̊JKvƂȂ邾낤Dʂ̃Av[`ƂāCtQ̂̍\pC̊𐧌ł悤ȒᕪqJ邱Ƃ\낤D
@̂悤ɁCtQ̃fՂƂtQ̐@\̉𖾂́CbƂĂ̏dvɂƂǂ܂炸C앨̐YɂďdvȌ`̉ǂɂȂ̂Ɗ҂D

<h2> </h2>
<ol>
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</ol>

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<span class='au'>cY,  o,  V </span>: <span class="ti">ԐztQƂ̎ê̍\݂͂ĂtQ@\̕q.</span> <span class='so'>wƐ, 50, 654-659 (2012)</span>
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<span class="author">  ViHiroyuki Tsujij</span>
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